Manhattan apparel designer Roxy Rae Vaughn, who's also a still-in-the-closet, country line dancing queen, is tired of being a Fifth Avenue up-and-comer. Against her parent’s wishes and without their financial backing, she moves to Nashville, Tennessee and opens a boutique in a local tractor supply store, the only retail space she can afford. Short on cash and luck, she rear-ends a pick-up truck belonging to a tomato-growin', bootscootin' cowboy who is anything but the perfect fit for her career plans, although he is her ideal dance partner. Can Roxy accept that her best fit is on the dance floor moving to her own style and her cowboy's lead? 

Tomato farmer by day and bootscootin' instructor by night Zayne McDonald doesn't give a damn about winning Nashville's heirloom tomato contest, even though his late father did. Zayne wants to honor his father, but what he really wants is to line dance his way into a winner's spotlight. When he and Roxy are discovered by a huge dance show producer, Zayne learns it's partnering with Roxy that's his winning hybrid mix. That is, if he can keep her and the tomato contest away from Beefsteak Jack Baudlin, the toughest tomato man in CMT country.